Warwick Davis
Master Moley (voice)
Gemma Arterton
Mona Lisa (voice)
Togo Igawa
Mystic Mole (voice)
Richard E. Grant
The Gardener (voice)
Charles Dance
Mr. James Moley (voice)
Julie Walters
Mrs. Elizabeth Moley / Queen (voice)
Calum Callaghan
Milo / American Elder (voice)
Amanda Drew
Melony (voice)
Sean Baker
Royal Guard / Guest (voice)
James Reatchlous
The Duke (voice)
Tony Nottage
Geeky Mole (voice)
Arthur Landon
Rodrigo (voice)
Leia Nottage
Child Mole (voice)
Rupert Holliday-Evans
(voice)