All Cast

Ray Romano

Ray Romano

Manny (voice)

John Leguizamo

John Leguizamo

Sid (voice)

Denis Leary

Denis Leary

Diego (voice)

Queen Latifah

Queen Latifah

Ellie (voice)

Seann William Scott

Seann William Scott

Crash (voice)

Josh Peck

Josh Peck

Eddie (voice)

Jay Leno

Jay Leno

Fast Tony (voice)

Will Arnett

Will Arnett

Lone Gunslinger Vulture (voice)

Chris Wedge

Chris Wedge

Scrat (voice)

Peter Ackerman

Peter Ackerman

Dung Beetle Dad (voice)

Caitlin Rose Anderson

Caitlin Rose Anderson

Glypto Boy Billy / Beaver Girl (voice)

Connor Anderson

Connor Anderson

Rhino Boy / Beaver Boy (voice)

Joseph Bologna

Joseph Bologna

Mr. Start (voice)

Jack Crocicchia

Jack Crocicchia

Elk Boy (voice)

Peter DeSève

Peter DeSève

Condor Chick (voice)

Ariel Winter

Ariel Winter

Additional Voices (voice)

Clea Lewis

Clea Lewis

Female Mini Sloth (voice)

Stephen Root

Stephen Root

Aardvark Dad (voice)

Nicole DeFelice

Nicole DeFelice

Start Girl (voice)

Debi Derryberry

Debi Derryberry

Diatryma Mom (voice)

Marshall Efron

Marshall Efron

Start Dad (voice)

Tom Fahn

Tom Fahn

Stu / Male Ox (voice)

Jason Fricchione

Jason Fricchione

Molehog Grandpa / Shovelmouth Male (voice)

James Edmund Godwin

James Edmund Godwin

Traffic Vulture (voice)

George Jacobs

George Jacobs

Beaver Dad (voice)

Brian Scott McFadden

Brian Scott McFadden

Glyptodon (voice)

Jansen Panettiere

Jansen Panettiere

Shovelmouth Boy (voice)

Gregory Romano

Gregory Romano

Start Boy 1 (voice)

Matt Romano

Matt Romano

Start Boy 2 (voice)

Carlos Saldanha

Carlos Saldanha

Dodo (voice)

Manoela Scarpa Saldanha

Manoela Scarpa Saldanha

Molehog Kid 1 (voice)

Sofia Scarpa Saldanha

Sofia Scarpa Saldanha

Molehog Kid 2 (voice)

Reyna Shaskan

Reyna Shaskan

Diego's Bird Girl (voice)

James Sie

James Sie

Freaky Male (voice)

Cindy Slattery

Cindy Slattery

Aardvark Mom (voice)

Mindy Sterling

Mindy Sterling

Female Ox (voice)

Renée Taylor

Renée Taylor

Mrs. Start (voice)

Alan Tudyk

Alan Tudyk

Cholly (voice)

Claudia Besso

Claudia Besso

Additional Voices (voice)

Madeleine Martin

Madeleine Martin

Additional Voices (voice)

Sherry Lynn

Sherry Lynn

Additional Voices (voice)

Mickie McGowan

Mickie McGowan

Additional Voices (voice)

Emily Anderson

Emily Anderson

Additional Voices (voice)

Jack Angel

Jack Angel

Additional Voices (voice)

Shane Baumel

Shane Baumel

Additional Voices (voice)

Bob Bergen

Bob Bergen

Additional Voices (voice)

Kyle J. Bogert

Kyle J. Bogert

Additional Voices (voice)

Rodger Bumpass

Rodger Bumpass

Additional Voices (voice)

Paul Butcher

Paul Butcher

Additional Voices (voice)

Blanca Camacho

Blanca Camacho

Additional Voices (voice)

John Cygan

John Cygan

Additional Voices (voice)

Jennifer Darling

Jennifer Darling

Additional Voices (voice)

Olivia DeLaurentis

Olivia DeLaurentis

Additional Voices (voice)

Harrison Fahn

Harrison Fahn

Additional Voices (voice)

Bill Farmer

Bill Farmer

Additional Voices (voice)

Crispin Freeman

Crispin Freeman

Additional Voices (voice)

Tim Kirkpatrick

Tim Kirkpatrick

Additional Voices (voice)

David H. Kramer

David H. Kramer

Additional Voices (voice)

Danny Mann

Danny Mann

Additional Voices (voice)

Laura Marano

Laura Marano

Additional Voices (voice)

Matt McCarthy

Matt McCarthy

Additional Voices (voice)

Laraine Newman

Laraine Newman

Additional Voices (voice)

Jan Rabson

Jan Rabson

Additional Voices (voice)

Zack Shada

Zack Shada

Additional Voices (voice)

Thomas Sharkey

Thomas Sharkey

Additional Voices (voice)

Ross Simanteris

Ross Simanteris

Additional Voices (voice)

Rachel Stand

Rachel Stand

Additional Voices (voice)

Spencer Lacey Ganus

Spencer Lacey Ganus

Additional Voices (voice)

Jess Harnell

Jess Harnell

Additional Voices (voice)

Alex Sullivan

Alex Sullivan

James (voice)